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SPECIALIZATION, SPECIATION, AND RADIATION This page intentionally left blank SPECIALIZATION, SPECIATION, AND RADIATION THE EVOLUTIONARY BIOLOGY OF HERBIVOROUS INSECTS Edited by KELLEY JEAN TILMON UNIVERSITY OF CALIFORNIA PRESS Berkeley Los Angeles London University of California Press, one of the most distinguished university presses in the United States, enriches lives around the world by advancing scholarship in the humanities, social sciences, and natural sciences. Its activities are supported by the UC Press Foundation and by philanthropic contributions from individuals and institutions. For more information, visit University of California Press Berkeley and Los Angeles, California University of California Press, Ltd. London, England© 2008 by the Regents of the University of California Library of Congress Cataloging-in-Publication Data Specialization, speciation, and radiation : the evolutionary biology of herbivorous insects / edited by Kelley Jean Tilmon. p. cm. Includes bibliographical references and index. ISBN 978-0-520-25132-8 (case : alk. paper) 1. Insects—Evolution. 2. Insects—Behavior. 3. Insect–plant relationships. I. Tilmon, Kelley Jean. QL468.7.S64 2008 595.7138—dc22 2007032362 Manufactured in the United States of America 10 09 08 10 9 8 7 6 5 4 3 2 1 The paper used in this publication meets the minimum requirements of ANSI/NISO Z39.48-1992 (R 1997) (Permanence of Paper). Cover photograph: A specialist caterpillar (Lepidoptera: Limacodidae) with brilliant warning coloration and stinging hairs feeds on a leaf of Macaranga tsonane (Euphorbiaceae) in the lowland rainforest of Papua New Guinea. Photo by George Weiblen. This book is dedicated to Tom Wood, whose abiding love of the Membracidae led him to study them in all their measure, and who was all that a naturalist, a mentor, and a friend should be. This page intentionally left blank ...