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Princeton University Press
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The most authoritative illustrated book on flying reptiles available

For 150 million years, the skies didn't belong to birds—they belonged to the pterosaurs. These flying reptiles, which include the pterodactyls, shared the world with the nonavian dinosaurs until their extinction 65 million years ago. Some pterosaurs, such as the giant azhdarchids, were the largest flying animals of all time, with wingspans exceeding thirty feet and standing heights comparable to modern giraffes. This richly illustrated book takes an unprecedented look at these astonishing creatures, presenting the latest findings on their anatomy, ecology, and extinction.

Pterosaurs features some 200 stunning illustrations, including original paintings by Mark Witton and photos of rarely seen fossils. After decades of mystery, paleontologists have finally begun to understand how pterosaurs are related to other reptiles, how they functioned as living animals, and, despite dwarfing all other flying animals, how they managed to become airborne. Here you can explore the fossil evidence of pterosaur behavior and ecology, learn about the skeletal and soft-tissue anatomy of pterosaurs, and consider the newest theories about their cryptic origins. This one-of-a-kind book covers the discovery history, paleobiogeography, anatomy, and behaviors of more than 130 species of pterosaur, and also discusses their demise at the end of the Mesozoic.


  • The most comprehensive book on pterosaurs ever published

  • Features some 200 illustrations, including original paintings by the author

  • Covers every known species and major group of pterosaurs

  • Describes pterosaur anatomy, ecology, behaviors, diversity, and more

  • Encourages further study with 500 references to primary pterosaur literature



Table of Contents

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  1. Cover
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  1. Title Page, Copyright, Dedication
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  1. Contents
  2. p. vii
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  1. Preface
  2. pp. ix-x
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  1. Acknowledgments
  2. p. xi
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  1. 1. Leathery-WingedHarpies
  2. pp. 1-3
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  1. 2. Understanding the Flying Reptiles
  2. pp. 4-11
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  1. 3. Pterosaur Beginnings
  2. pp. 12-22
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  1. 4. The Pterosaur Skeleton
  2. pp. 23-38
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  1. 5. Soft Bits
  2. pp. 39-55
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  1. 6. Flying Reptiles
  2. pp. 56-63
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  1. 7. Down from the Skies
  2. pp. 64-73
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  1. 8. The Private Lives of Pterosaurs
  2. pp. 74-89
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  1. 9. The Diversity of Pterosaurs
  2. pp. 90-94
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  1. 10. Early Pterosaurs and Dimorphodontidae
  2. pp. 95-103
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  1. 11. Anurognathidae
  2. pp. 104-112
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  1. 12. “Campylognathoidids”
  2. pp. 113-122
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  1. 13. Rhamphorhynchidae
  2. pp. 123-134
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  1. 14. Wukongopteridae
  2. pp. 135-142
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  1. 15. Istiodactylidae
  2. pp. 143-151
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  1. 16. Ornithocheiridae
  2. pp. 152-163
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  1. 17. Boreopteridae
  2. pp. 164-169
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  1. 18. Pteranodontia
  2. pp. 170-182
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  1. 19. Ctenochasmatoidea
  2. pp. 183-200
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  1. 20. Dsungaripteroidea
  2. pp. 201-210
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  1. 21. Lonchodectidae
  2. pp. 211-215
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  1. 22. Tapejaridae
  2. pp. 216-227
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  1. 23. Chaoyangopteridae
  2. pp. 228-233
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  1. 24. Thalassodromidae
  2. pp. 234-243
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  1. 25. Azhdarchidae
  2. pp. 244-258
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  1. 26. The Rise and Fall of the Pterosaur Empire
  2. pp. 259-264
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  1. References
  2. pp. 265-282
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  1. Index
  2. pp. 283-292
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