New Perspectives on Horned Dinosaurs
The Royal Tyrrell Museum Ceratopsian Symposium
Publication Year: 2010
Easily distinguished by the horns and frills on their skulls, ceratopsians were one of the most successful of all dinosaurs. This volume presents a broad range of cutting-edge research on the functional biology, behavior, systematics, paleoecology, and paleogeography of the horned dinosaurs, and includes descriptions of newly identified species.
Published by: Indiana University Press
Download PDF (81.8 KB)
Download PDF (78.5 KB)
...that Phil is now at the University of Alberta); (2) Michael Ryancated to ceratopsians at the Tyrrell Museum, we realized that itirrefutable ‘‘king’’ of ceratopsian dinosaurs, Peter Dodson. Pe-saurs, and here he reflects on all he has learned over the years,while providing historical perspective about those he has ‘‘in-...
Download PDF (31.2 KB)
Andy Farke for his diligence and efforts in cheerfully and effec-huge contribution in serving as Editorial Assistant for this proj-...
List of Contributors
Download PDF (52.2 KB)
List of Reviewers
Download PDF (98.0 KB)
PART ONE § OVERVIEW
Download PDF (62.9 KB)
1. Forty Years of Ceratophilia
Download PDF (336.6 KB)
...or at least the senior citizen, of ceratopsian studies. Of course‘‘stuff.’’ A vivid early encounter with dinosaurs came when mymother, a lover of classical music, took me to see Walt Disney’sstarted with the beginning of unicellular life in the primordialstirring chords of Stravinsky’s Sacre du Printemps. Growing up...
PART TWO § SYSTEMATICS AND NEW CERATOPSIANS
Download PDF (62.9 KB)
2. Taxonomy, Cranial Morphology, and Relationships of Parrot-Beaked Dinosaurs(Ceratopsia: Psittacosaurus)
Download PDF (1.1 MB)
...lished, including the description of two new species (P. meiley-species of Psittacosaurus, P. lujiatunensis (Zhou et al. 2006b) andFIGURE 2.1. Paleogeographic distribution of psittacosaurids and location of particularly fossiliferous orsingular localities. Locality abbreviations, 1: Shestakovo (Siberia); 2: Delunshan (Xinjiang); 3: Bulasu-tuin; 4: Ondai Sair; 5: Khobur; 6: Oshih; 7: Ulan Osh; 8: Khuren Dukh; 9: Sharalin Ula; 10: Tsakurt;...
3. A New Species of Archaeoceratops (Dinosauria: Neoceratopsia) from the Early Cretaceous of the Mazongshan Area, Northwestern China
Download PDF (279.9 KB)
A New Species of Archaeoceratops (Dinosauria: Neoceratopsia) from therioide, has been studied (Li et al. 2007). Here, we describe a newa partial skull extending from the right premaxilla to jugal andEtymology. ‘‘Yujingzi’’ (Chinese): name of the basin wherethat do not exist in A. oshimai: a laterally deflected rostral end...
4. A Redescription of the Montanoceratops cerorhynchus Holotype with a Review of Referred Material
Download PDF (261.1 KB)
...‘‘Leptoceratops’’ cerorhynchus (Brown and Schlaikjer 1942) was‘‘protoceratopsid’’ Leptoceratops. Shortly before its description,genus Montanoceratops for the type of ‘’’Leptoceratops’’ cero-ska 1975; Ostrom 1966), a fact also reflected in several cladisticceratops cerorhynchus as the sister taxon to Ceratopsidae (Chin-...
5. First Basal Neoceratopsian from the Oldman Formation (Belly River Group), Southern Alberta
Download PDF (239.5 KB)
TETSUTO MIYASHITA, PHILIP J. CURRIE, AND BRENDA J. CHINNERY-ALLGEIERmation (a rationale for this interpretation in ‘‘Discussion’’).to the interfrontal suture; straight, transverse postorbital ridge;TMP 87.89.8 (Fig. 5.3) is an isolated right frontal. The anteriorber 61, Fig. 5.4A). This is consistent with an allometric trend in...
6. Zuniceratops christopheri: The North American Ceratopsid Sister Taxon Reconstructed on the Basis of New Data
Download PDF (255.6 KB)
...ered as floated, but associated, skeletal fragments in 1996. Pub-lac: lacrimal; lsp: laterosphenoid; mjc: maxillary-jugal contactsurface; mx: maxilla in left lateral view; n: naris opening; na:nasal in left lateral view; oc: occipital condyle; pa1: parietaltary in left lateral view; pmx: right premaxilla in medial view;...
7. Horned Dinosaurs (Ornithischia: Ceratopsidae) from the Upper Cretaceous (Campanian)Cerro del Pueblo Formation, Coahuila, Mexico
Download PDF (461.8 KB)
MARK A. LOEWEN, SCOTT D. SAMPSON, ERIC K. LUND, ANDREW A. FARKE,RUBÉN A. RODRÍGUEZ-DE LA ROSA, MICHAEL A. GETTY, AND DAVID A. EBERTH100 loewen, sampson, lund, farke, aguillón-martínez, de leon, rodríguez-de la rosa, getty, and eberthC. russelli Sternberg 1940, and C. irvinensis Holmes et al. 2001.Soegaard et al. 1997; Kirkland et al. 2000; Eberth et al. 2004)....
8. New Basal Centrosaurine Ceratopsian Skulls from the Wahweap Formation (Middle Campanian), Grand Staircase–Escalante National Monument, Southern Utah
Download PDF (797.2 KB)
...is a smaller ‘‘epinasal’’; (3) a large accessory antorbitalcally critical cranial remains have yet to be identified in thesenasal area and the rear of the frill, we concluded that given thecomplete skull lying on its left side; part of the right side hadsian skulls. Both skulls are curated into the vertebrate paleon-...
9. A New Pachyrhinosaurus-Like Ceratopsid from the Upper Dinosaur Park Formation (Late Campanian) of Southern Alberta, Canada
Download PDF (432.6 KB)
...sp. is known from Alaska (Fiorillo et al. this volume). In regions‘‘pachyrhinosaurs’’ (Pachyrhinosaurus + Achelousaurus) distin-while usually described as being a ‘‘boss,’’ is more typically aFIGURE 9.1. Locality map. TMP 2002.76.1 occurs at the southeastern margin ofDinosaur Provincal Park, Alberta. Inset shows the location (star) relative to Alberta....
10. New Material of ‘‘Styracosaurus’’ ovatus from the Two Medicine Formation of Montana
Download PDF (373.8 KB)
...new generic name, Rubeosaurus, is created for ‘‘S.’’ ovatus,centrosaurine ceratopsids: ‘‘Brachyceratops montanensis’’ (Gil-lated partial specimens, respectively. The material of ‘‘Brachy-ceratops’’ includes the holotype partial juvenile skull, USNMuals excavated from the same quarry (Gilmore 1917). ‘‘Brachy-...
11. A New Chasmosaurine (Ceratopsidae, Dinosauria) from the Upper Cretaceous OjoAlamo Formation (Naashoibito Member), San Juan Basin, New Mexico
Download PDF (372.7 KB)
...with ‘‘deep vascular grooves’’ and a small, smooth ‘‘supraorbi-the Kirtland Formation (‘‘upper shale’’ of traditional usage)light brown-tan preservation of the ‘‘supraorbital horncore’’with ‘‘deep radiating vascular impressions,’’ a third consistingof ‘‘similar fragments,’’ and the fourth consisting of isolated...
12. A New Chasmosaurine Ceratopsid from the Judith River Formation, Montana
Download PDF (260.7 KB)
...genera in two clades (‘‘subfamilies’’), Centrosaurinae andclonius crassus, M. fissus, M. recurvicornus, and M. sphenocerus;referred to as the ‘‘Mansfield Bonebed’’ in reference to thester in Greek mythology with ‘‘hair’’ comprised of snakes andlarge, thick snake-like spikes that extend from the lateral mar-...
13. Description of a Complete and Fully Articulated Chasmosaurine Postcranium Previously Assigned to Anchiceratops (Dinosauria: Ceratopsia)
Download PDF (326.2 KB)
...of the type of A. ‘‘longirostris’’ (Sternberg 1929) in place.detail, paying particular attention to those defining features ofcranial skeleton with associated frill fragments. The ‘‘anteriorportion of the beak’’ reported by Sternberg in his field notesthird digits of the left manus depicted in both old field photo-...
14. A New, Small Ceratopsian Dinosaur from the Latest Cretaceous Hell Creek Formation, Northwest South Dakota, United States: A Preliminary Description
Download PDF (561.7 KB)
A New, Small Ceratopsian Dinosaur from the Latest Cretaceous Helltion is currently represented by four taxa: Triceratops horridusproportions of Triceratops horridus (adult skull length approxi-FIGURE 14.1. Holotype skull of Tatankaceratops sacrisonorum (BI-6226). (A) Known cranial material; (B) reconstruction of the skull ofTatankaceratops sacrisonorum (BHI-6226) superimposed on the reconstructed skull of Triceratops horridus (TCM 2001.93.1; in grey)....
PART THREE § ANATOMY, FUNCTIONAL BIOLOGY, AND BEHAVIOR
Download PDF (62.9 KB)
15. Comments on the Basicranium and Palate of Basal Ceratopsians
Download PDF (378.5 KB)
...(Zhao et al. 1999); Liaoceratops (Xu et al. 2002); Hongshanosau-et al. 2006), all from China. In recent years, there has also beention of new species: P. xinjiangensis (Sereno and Chao 1988); P.meileyingensis (Sereno et al. 1988); P. neimongoliensis (RussellGansu; P. sibiricus from Siberia (Averianov et al. 2006); P. lu-...
16. Mandibular Anatomy in Basal Ceratopsia
Download PDF (295.7 KB)
...observation of the skulls of basal ceratopsians is necessary. Un-straight with very little change in width except for at the cau-circular as in Psittacosaurus (Fig. 16.1A). Unlike in Psittacosau-slightly in lateral view. The preserved rostral tip is a few milli-angular, if not higher. This feature is similar to that of Yinlong...
17. Histological Evaluation of Ontogenetic Bone Surface Texture Changes in the Frill of Centrosaurus apertus
Download PDF (462.2 KB)
...absolute age data and soft tissue anatomy is a critical issue inadult, and adult, as applied to fossil vertebrates, generally im-This is in sharp contrast to the use of the terms juvenile, sub-1997); the reverse is true in extant birds (Craighead and Stock-Deratzian et al. 2007) have addressed the possibility of predict-...
18. Modeling Structural Properties of the Frill of Triceratops
Download PDF (163.2 KB)
...relatively shorter frill (as compared to basal skull length) thanters has led to speculation that the frill of Triceratops was analso noted that the frill in some ceratopsids was constructed asa ‘‘perfect frame,’’ and hypothesized that the parietal fenestraeloads. Here, we investigate the structural properties of the un-...
Download PDF (3.0 MB)
Pl a t e 1. (Figure 2.2) Psittacosaurus sp. (LHPV2). Skull in (A) lateral-view; (B) enlarged view of postorbital-jugal horn in lateral view; Pl a t e 2. (Figure 2.4) Psittacosaurus major (LH PV1). (A) Stereopairs of anterior palate in ventral view; (B) stereopairs of posterior palate and braincase in posterioventral view. Pl a t e 5. (Figure 2.18) Psittacosaurus mongoliensis (AMNH 6535) stereopairs of hatchling cranium in ...
19. New Evidence Regarding the Structure and Function of the Horns in Triceratops (Dinosauria: Ceratopsidae)
Download PDF (421.5 KB)
...(Gilmore 1920). Hatcher et al. (1907) attributed the black layercovering of a Triceratops horn that may relate to Hatcher’s orig-FIGURE 19.1. SUP 9713.0 in the quarry site before removal. Skull is upside down and articulated except for supraorbital horns andone epijugal. Left supraorbital horn removed and not shown. Scale bar is 10 cm.sinus matrix; nh: nasal horn; r: rostral; sh: right supraorbital...
20. Evolutionary Interactions between Horn and Frill Morphology in Chasmosaurine Ceratopsians
Download PDF (197.1 KB)
...reptiles, or been used as a display structure to attract a mate.(1928) suggested that the large, solid frill of Triceratops couldEvolutionary Interactions between Horn and Frill Morphology in Chasmosaurine Ceratopsians 283Farke 2007). However, frills with large parietal fenestrae wouldvery little resistance to torsional stresses caused by a predatory...
21. Skull Shapes as Indicators of Niche Partitioning by Sympatric Chasmosaurine and Centrosaurine Dinosaurs
Download PDF (306.2 KB)
...sus diurnal feeding times of birds of prey such as owls (Strigi-between species, such as the differences in body size and denti-tions of sympatric carnivores, which are associated with differ-have also been observed in the fossil record of vertebrates that2005). Similar stratigraphic positions for sets of fossils imply...
22. The Function of Large Eyes in Protoceratops: A Nocturnal Ceratopsian?
Download PDF (406.4 KB)
The Function of Large Eyes in Protoceratops: A Nocturnal Ceratopsian?Few vertebrates, either fossil or extant, are as well-represented1922, the first fossils of Protoceratops andrewsi were recoveredAsiatic Expedition collected a dozen skeletons of Protoceratops,FIGURE 22.1. Skull of Protoceratops andrewsi (AMNH 6466) and comparisons with Centrosaurus apertus. (A) A large ?female showing the...
23. A Semi-Aquatic Life Habit for Psittacosaurus
Download PDF (268.7 KB)
...‘‘kick.’’ Other morphological features that support an in-(5) ‘‘bristle-like’’ integumentary structure on the dorsalstrictly terrestrial forms (Bakker 1986). Not only did the rein-swamps, rivers and lakes, but it also showed that they weren’tnecessarily sluggish reptiles. In the last few decades, however,...
24. Habitual Locomotor Behavior Inferred from Manual Pathology in Two Late CretaceousChasmosaurine Ceratopsid Dinosaurs, Chasmosaurus irvinensis (CMN 41357) and Chasmosaurus belli(ROM 843)
Download PDF (320.2 KB)
Chasmosaurus irvinensis (CMN 41357) and Chasmosaurus belli (ROM 843)Paleopathological ‘‘diagnosis’’ is necessarily an approximateare not attempting to ‘‘cure’’ fossils—instead, they are occu-et al. 1994). Moreover, strict diagnosis of disease in the past is(Buikstra 1981). Assertion of diagnostic authority or uncritical...
25. Paleopathologies in Albertan Ceratopsids and Their Behavioral Significance
Download PDF (830.9 KB)
...the occurrences of injuries (i.e., avulsions, fractures, healinglittle detailed attention in the distant past. Most early reportsing the evolution of ‘‘freak’’ features such as frills found in‘‘. . . pathological (fungal?) corrosion and without geneticstress fracture in a ceratopsian phalanx; the first paper focused...
PART FOUR § HORNED DINOSAURS IN TIME AND SPACE
Download PDF (78.7 KB)
26. An Update on the Paleobiogeography of Ceratopsian Dinosaurs
Download PDF (421.1 KB)
The term ‘‘paleobiogeography’’ means, in essence, the studyveal aspects of its evolutionary history not readily ascertainedpothesis (i.e., Sereno 1999, 2000). However, in the last 10–15the ancient globe at various time periods, lost data from inevi-differences can, in fact, significantly alter interpretations of...
27. Unraveling a Radiation: A Review of the Diversity, Stratigraphic Distribution, Biogeography, and Evolution of Horned Dinosaurs (Ornithischia: Ceratopsidae)
Download PDF (526.3 KB)
Unraveling a Radiation: A Review of the Diversity, Stratigraphicdivided into two subclades (‘‘subfamilies’’): Centrosaurinae,highly adorned frills (e.g., Styracosaurus albertensis); and Chas-skull materials); (2) putative species-specific signaling struc-discoveries that greatly increase the material basis of the clade....
28. A Review of Ceratopsian Paleoenvironmental Associations and Taphonomy
Download PDF (169.5 KB)
A Review of Ceratopsian Paleoenvironmental Associations and Taphonomy 429potential significance of these patterns in the context of long-land; Eberth et al.; Fiorillo et al.; Ford and Martin; Getty et al.;tion with the patterns and data-gaps identified here, will stim-logical associations, adaptability, and preservational processes...
29. Behavioral Interpretations from Ceratopsid Bonebeds
Download PDF (131.2 KB)
...stitute evidence of gregarious, herding behavior. There is littleand to a lesser extent, the scientific literature (Currie and Dod-clade? Is there any evidence in the fossil record for behavioralfer quite sharply in their relative abundance in the rock recordpaleochannel (here termed ‘‘higher energy’’) and overbank...
30. Paleontology and Paleoenvironmental Interpretation of the Kikak-Tegoseak Quarry (Prince Creek Formation: Late Cretaceous), Northern Alaska: A Multi-Disciplinary Study of a High-Latitude Ceratopsian Dinosaur Bonebed
Download PDF (586.3 KB)
Kikak-Tegoseak Quarry (Prince Creek Formation: Late Cretaceous),ANTHONY R. FIORILLO, PAUL J. MCCARTHY, PETER P. FLAIG, ERIK BRANDLEN,DAVID W. NORTON, PIERRE ZIPPI, LOUIS JACOBS, AND ROLAND A. GANGLOFFPaleontology and Paleoenvironmental Interpretation of the Kikak-Tegoseak Quarry 457to gain similar insights into ancient terrestrial ecosystems and...
31. Taphonomy of Horned Dinosaurs (Ornithischia: Ceratopsidae) from the Late Campanian Kaiparowits Formation, Grand Staircase–Escalante National Monument, Utah
Download PDF (317.2 KB)
Taphonomy of Horned Dinosaurs from the Late Campanian Kaiparowits Formation 4792007; Gates et al. in press; Zanno et al. in press). By far the mostter refine the stratigraphic framework for the target formationslogic research relevant to the region’s vertebrate paleontologyparowits Formation thus correlates with the fossiliferous inter-...
32. A Centrosaurine Mega-Bonebed from the Upper Cretaceous of Southern Alberta: Implications for Behavior and Death Events
Download PDF (358.1 KB)
...badlands at DPP significantly limits one’s ability to trace in-‘‘Hilda,’’ 80 km to the southeast along the South Saskatche-(H97-02, 04, 06, 08, 11) are in bold type. Shaded area indicateshost bed (see Figs. 32.3 and 32.4). Section, township, and rangeA Centrosaurine Mega-Bonebed from the Upper Cretaceous of Southern Alberta 497...
33. Insect Trace Fossils Associated with Protoceratops Carcasses in the Djadokhta Formation (Upper Cretaceous), Mongolia
Download PDF (470.4 KB)
Insect Trace Fossils Associated with Protoceratops Carcasses in theet al. 1993; Fastovsky et al. 1997; Hasiotis et al. 1999; Paik 2000;surrounding regions, showing localities where Protoceratops havebeen recovered. (A) Inset of central Asia; (B) topographic map ofwere, from that point, left in association with the skeleton. The...
34. Faunal Composition and Significance of High-Diversity, Mixed Bonebeds Containing Agujaceratops mariscalensis and Other Dinosaurs, Aguja Formation (Upper Cretaceous), Big Bend, Texas
Download PDF (403.8 KB)
Faunal Composition and Significance of High-Diversity, Mixed Bonebedssaur remains, as well as rich and diverse vertebrate microfossilSignificance of High-Diversity, Mixed Bonebeds Containing Agujaceratops mariscalensis 521in the Tornillo Basin of west Texas during the Late Cretaceous–34.2; Nordt et al. 2003; Atchley et al. 2004). The A. mariscalen-...
PART FIVE § HISTORY OF HORNED DINOSAUR COLLECTION
Download PDF (62.9 KB)
35. Lost in Plain Sight: Rediscovery of William E. Cutler’s Missing Eoceratops
Download PDF (208.2 KB)
Lost in Plain Sight: Rediscovery of William E. Cutler’s Missing Eoceratopsa problem to society or especially to their families. Often they werehave them safely far away for good, wealthier families enticed thefor a boat ride overseas and a sum of money to get started. To keepthem away, a ‘‘remittance check’’ or allowance was sent to them at...
36. Historical Collecting Bias and the Fossil Record of Triceratops in Montana
Download PDF (780.3 KB)
By anyone’s measure, Triceratops, or ‘‘three-horned face,’’ isthe Laramie ‘‘Ceratops’’ beds, now formally referred to as theinitiated the Hell Creek Project, at the time a five-year paleon-Historical Collecting Bias and the Fossil Record of Triceratops in Montana 553most part of the section; and (2) these exposures are relatively...
Download PDF (86.1 KB)
...number of authors in this book. In fact, it was very rare duringthe first century of ceratopsian studies (starting with Cope’sLull’s ‘‘Review of the Ceratopsia’’ in 1933. Several other cera-John B. Hatcher was even represented ‘‘in the flesh’’ by one ofwho is still an undergraduate at the time this was written)—are...
Download PDF (297.8 KB)
Page Count: 656
Publication Year: 2010
Series Title: Life of the Past