Cover

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Title Page, Series Page, Copyright

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pp. i-iv

Contents

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pp. v-vi

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Preface

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pp. vii-viii

...containing just recognizably different eggs, that prompted a much more thorough look: it appeared that contrary to what natural selection usually dictates, a female ostrich seemed to be doing all of another female's work for her. At that time...

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1 The Ostrich

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pp. 1-18

...five to only two in the ostrich is probably another adaptation for fast running. The large inner toe (originally the third) and occasionally the smaller outer toe (originally the fourth) carry a powerful...

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2 Methods

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pp. 19-32

...chapters, a summary is first provided in this chapter of the environment in which ostriches were studied in Kenya. The methods by which the birds could be observed and recognized individually are outlined, and the kinds of observations that were...

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3 The Population

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pp. 33-48

...distinguishable and the proportions of the population falling into those age and sex categories. The relative advantages of different reproductive strategies depend partly on the sex ratio of breeding birds within the population; biases due to differential conspicuousness...

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4 The Breeding System

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pp. 49-70

...more detailed analyses of the reproductive options open to different categories of breeding birds. This chapter summarizes when and where ostrich nests in general were started, and by whom; how they grew; how they were attended; what other birds laid in them and when; how large they finally ended up...

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5 Ecological Aspects

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pp. 71-101

...these, two of the most prevalent costs are the loss of eggs and the death of birds, and in this chapter I endeavour to quantify some of the effects of the natural hazards that ostriches face in Tsavo West National Park. Predation on nests...

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6 Strategies Adopted by Major Females

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pp. 102-123

...or minor role, and the relative reproductive pay-offs of the two different strategies, are considered in Chapter 7. In this chapter, for major females already having begun a nest, I will consider the options open to them at various stages, and the...

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7 Major and Minor Female Strategies

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pp. 124-137

...a result, it is becoming clear why, if she is a major female, an ostrich hen behaves as she does. We must now consider what a minor female's options are. I examine to what extent the minor hen strategy was consistent and whether there were different...

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8 Male Strategies

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pp. 138-158

...he goes out of breeding condition at incubation, whereas if he does not have a nest, he remains in breeding condition for longer and continues to mate promiscuously with females who lay in others' nests. In this chapter, I consider first what proportion...

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9 Discussion: The Evolution and Maintenance of the Communal Nesting System

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pp. 159-187

...pressures are operating within the ostrich communal nesting system to influence the choices that individual birds make at different stages of their nesting. It is the consequences of those choices that natural selection acts upon: selection will...

References

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pp. 188-192

Subject Index

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pp. 193-197