Cover

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pp. 1-1

Title Page, Copyright

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pp. 2-5

Contents

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pp. v-viii

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Abstract

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pp. ix-x

Fifty-nine species, including 25 new species, of the marine calanoid family Heterorhabdidae are described and their geographic ranges defined from specimens found in midwater trawl and plankton net samples collected throughout the Atlantic, Pacific, and Indian Oceans. ...

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Acknowledgments

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pp. xi-xii

This study is based mainly on midwater trawl and plankton net samples housed in the Planktonic Invertebrates Collections at Scripps Institution of Oceanography. I wish to thank M. Ohman, curator, and A. Townsend, museum scientist, of these collections, who made the samples available for my study. ...

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Introduction

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pp. 1-2

The family Heterorhabdidae was erected by Sars (1902) to accommodate the genera Heterorhabdus Giesbrecht 1898 (published in Giesbrecht and Schmeil 1898), Haloptilus Giesbrecht 1898, and Augaptilus Giesbrecht 1889. Sars (1905) added Disseta Giesbrecht 1889 and a new genus, Mesorhabdus, ...

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Materials and Methods

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pp. 3-4

Most of the specimens examined in this study were picked from Isaacs-Kidd rnidwater trawl (IKMT) samples selected from the collections available at Scripps Institution of Oceanography (SIO). Additional specimens were obtained from IKMT, plankton net and MOCNESS (Multiple Opening1 Closing Net and Environmental Sensing System) ...

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Family Heterorhabdidae Sars 1902

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pp. 5-10

Diagnosis. 1) Left caudal ramus normally fused to anal segment, longer than right ramus. 2) Fourth marginal seta (counted from lateral) of left caudal ramus naked and greatly elongated. 3) Basis of male right P5 with a large, plumose inner lobe, which in some taxa is extremely large. ...

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Genus Disseta Giesbrecht 1889

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pp. 11-22

Diagnosis. All principal marginal setae of caudal rami except for 4th on left ramus armed with small spines in addition to normal setules. First segment of A1 with 1 3 3 3 setaelaesthetes (Fig.1-0. In antenna, all 8 exopodal segments each with a well-developed inner marginal seta (Fig.1-h). In mandible, masticatory edge with a basal spine, ...

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Genus Mesorhabdus Sars 1905

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pp. 23-36

Diagnosis. Third marginal seta of left caudal ramus and 3rd and 4th marginal setae of right caudal ramus armed with small spines in addition to normal setules (Fig.11-k). First segment of A1 with 1, 3, 3, 3 setae/aesthetes (Fig.10-g). In A2, 1st exopodal segment without seta and 2nd with a small marginal seta (Fig. 10-h). ...

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Genus Heterostylites Sars 1920

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pp. 37-50

Diagnosis. None of marginal setae of caudal rami armed with conspicuous spines in addition to normal setules. First segment of A1 (Fig.18-h) with 4 lobes bearing 1, 3, 2, 2 setaelaesthetes, respectively. In exopod of A2 (Fig.18-j), first 2 exopodal segments without setae. In mandible (Fig.19- a), masticatory edge with a basal spine and 3 or 4 long spiniform teeth, ...

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Genus Hemirhabdus Wolfenden 1911

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pp. 51-56

Diagnosis. Body massive. First segment of A1 (Fig.30-f) with 1, 3, 3, 3 setae/aesthetes. First and 2nd exopodal segments of A2 (Fig.30-h) each with a small inner marginal seta. Basis and 1st endopodal segment of Md with 1 and 2 setae, respectively (Fig.30-i); 2nd endopodal segment without appendicular setae; ...

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Genus Neorhabdus Heptner 1972

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pp. 57-70

Diagnosis. Body massive. In 1st segment of A1 (Fig.34-h), 1st lobe with 1 seta, 2nd with 2 setae and 1 aesthete, 3rd with 2 setae and 1 or more aesthetes, and 4th with 2 setae and 2 or more aesthetes. First and 2nd exopodal segments of A2 without setae (Fig.34-i); inner and outer lobes of 2nd endopodal segment without appendicular setae. ...

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Genus Paraheterorhabdus Brodsky 1950

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pp. 71-88

Diagnosis. Dorsally, forehead (Fig.43-e) with a midanterior tubercular process that continues into rostra1 base. Third marginal seta of left caudal ramus and 3rd and 4th marginal setae of right caudal ramus armed with spines in addition to normal setules. In 1st segment of A1 (Fig.43-h), 1st lobe with 1 seta, 2nd to 4th each with 2 setae and 1 aesthete. ...

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Genus Heterorhabdus Giesbrecht 1898

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pp. 89-140

Diagnosis. Dorsally, forehead (Fig.59-e) with a midanterior tubercular process, which extends ventrad into rostrum. Sternite of 1st pedigerous somite (Fig.59-a) with a spiny papilla. Left caudal ramus (Fig.59-c) fused with anal segment and distinctly longer than right. In 1st segment of A1 (Fig.59-j), 1st lobe with a small seta, ...

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Phylogeny

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pp. 141-142

As members of the superfamily Arietelloidea, the Heterorhabdidae share the general morphology of the body and appendages with the other members of the superfamily, in particular with the Lucicutiidae and Augaptilidae. The Heterorhabdidae can readily be distinguished from them and any other calanoid families by the following synapomorphies: ...

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Distribution

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pp. 143-146

All of the genera, subgenera, and species groups of the Heterorhabdidae defined in this study are circumglobal in distribution, occurring widely in all three major oceans. Disseta is a bathypelagic genus comprising 3 species, of which D. palumbii is a common circumglobal species occurring widely in the low and midlatitudes of all three major oceans. ...

Literature Cited

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pp. 147-150

Figures

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pp. 151-270

Back Cover

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pp. 284-284